Title: Classification of Vegetation Communities of Maryland

Classification of Vegetation Communities of Maryland: First Iteration

A Subset of the International Classification of Ecological
Communities: Terrestrial Vegetation of the United States
March 2004

V. HERBACEOUS VEGETATION
V.C.2.N.a. Permanently flooded temperate or subpolar hydromorphic rooted vegetation
V.C.2.N.a.102. NYMPHAEA ODORATA - NUPHAR SPP. PERMANENTLY FLOODED TEMPERATE HERBACEOUS ALLIANCE

White Water-lily - Yellow Pond-lily species Permanently Flooded Temperate Herbaceous Alliance


Concept: This alliance, common throughout most of the eastern and central United States and adjacent Canadian provinces, contains vegetation which may occur in a variety of slow-moving water bodies, including rivers, millponds, blackwater rivers, streams, shallow ponds or lakes, or on shores of deeper water bodies including freshwater tidal areas. The water depth is generally greater than 0.5 m and up to 2 m. Stands are dominated by hydromorphic rooted aquatic plants, typically Nuphar lutea (any of its various subspecies), with or without Nymphaea odorata. Emergent vegetation is less than 25%, and typically plant species diversity is low. Other species present may include Utricularia spp., Potamogeton spp., and others. In the north, Brasenia schreberi may be locally dominant. Other characteristic northern species include Nymphaea tetragona and Potamogeton amplifolius. Associates found in the Midwest include Polygonum amphibium. In the Southeast, examples may include the floating or emergent 'pad-leaved' species Nelumbo lutea or Nymphoides aquatica. Submerged aquatic species which may be present include Cabomba caroliniana, Ceratophyllum demersum, and Heteranthera dubia. Stands of this alliance are permanently to semipermanently flooded.

Comments: Field guidelines for separating floating-leaved aquatic alliances from submerged aquatic alliances are also needed.

Range: This alliance is common throughout most of the eastern and central United States and adjacent Canadian provinces. It is also found in Oregon, Washington, California, Idaho, Colorado, and possibly Wyoming (?).

States/Provinces: AL AR BC CA CO CT DE FL GA IA ID IL IN KY LA MA MB MD ME MI MN MO MS NC NH NJ NY OH OK ON OR PA RI SC TN TX VA VT WA WI WV WY?

TNC Ecoregions: 10:C, 20:C, 2:C, 31:C, 32:P, 36:C, 37:C, 39:C, 40:P, 41:C, 42:C, 43:C, 44:C, 45:C, 46:C, 47:C, 48:C, 49:C, 50:C, 51:C, 52:C, 53:C, 55:C, 56:C, 57:C, 58:C, 59:C, 60:C, 61:C, 62:C, 63:C, 6:C

USFS Ecoregions: 212Aa:CCC, 212Ba:CCC, 212Bb:CCC, 212Ca:CCC, 212Cb:CCC, 212Da:CCC, 212Db:CCP, 212Dc:CCC, 212Fa:CCP, 212Fb:CCP, 212Fc:CCP, 212Fd:CCP, 212Ga:CCP, 212Gb:CCP, 212Ha:CPP, 212Hb:CPP, 212He:CPP, 212Hh:CPP, 212Hi:CPP, 212Hj:CPP, 212Hk:CPP, 212Hl:CPP, 212Hm:CPP, 212Hn:CPP, 212Ho:CPP, 212Hp:CPP, 212Hq:CPP, 212Hr:CPP, 212Hs:CPP, 212Ht:CPP, 212Hu:CPP, 212Hv:CPP, 212Hw:CPP, 212Hx:CPP, 212Hy:CPP, 212Ib:CPP, 212Ja:CCP, 212Jb:CCP, 212Jc:CCP, 212Je:CCP, 212Jf:CCP, 212Jj:CCP, 212Jk:CCP, 212Jl:CCP, 212Jm:CCC, 212Jn:CCP, 212Jo:CCP, 212Jr:CCP, 212Ka:CPP, 212La:CCP, 212Lb:CCC, 212Lc:CCP, 212Ld:CCC, 212Ma:CPP, 212Mb:CPP, 212Na:CPP, 212Nb:CPP, 212Nc:CPP, 221Ah:CCC, 221Ai:CCC, 221Al:CCC, 221Am:CCC, 221Bd:CCP, 221Da:CCC, 221Db:CCC, 221Ea:CCC, 221Ed:CC?, 221Ef:CCC, 221He:CCC, 222Ch:CCC, 222Db:CCC, 222Gc:C??, 222Ha:CCC, 222Ja:CCC, 222Jb:CCC, 222Je:CCC, 222Ji:CCC, 222Jj:CCC, 222Kf:CCC, 222Kg:CCC, 222Kh:CCC, 222Kj:CCC, 231Bc:CCC, 231Ga:CCC, 231Gb:CCC, 231Gc:CCC, 232Ac:CCC, 232Bf:CCC, 232Bg:CCC, 232Bj:CC?, 232Ca:CCC, 232Cb:CCC, 232Cc:CC?, 232Cd:CCC, 232Ch:CCC, 232Dc:CCC, 234Ac:CC?, 234An:CCC, 242A:CC, 251Cf:CCC, 251Dd:CCC, 251Dg:CCC, 251Eb:CCC, M212Af:CCC, M212Bc:CCC, M212Bd:CCC, M212Cb:CCC, M212Cc:CCC, M212Ea:CCP, M212Eb:CCP, M221Aa:CCC, M221Ab:CCC, M221Ac:CCC, M221Ad:CCC, M221Ba:CCC, M221Bb:CCC, M221Bc:CCC, M221Bd:CCC, M221Be:CCC, M221Bf:CCC, M221Da:CCC, M221Dc:CCC, M221Dd:CCC, M331A:CC, M331D:CC, M331H:CC, M332E:CC, M333A:CC, M333B:CC, M333D:CC

Federal Lands: DOD (Eglin, Fort Benning); NPS (Acadia, Carl Sandburg Home, Isle Royale, Voyageurs); USFS (Angelina, Conecuh, Croatan?, Davy Crockett, Kisatchie, Ocala, Ozark, Sabine NF, Sam Houston?, Talladega); USFWS (Okefenokee, Reelfoot)

Synonymy: Spring-run Stream, in part (FNAI 1990); Open water/aquatic bed veg., natural impoundment pond (Ambrose 1990a); Small Depression Pond, in part (Schafale and Weakley 1990); Nuphar lutea herbaceous alliance, in part (Hoagland 1998a); L5D2aI1a. Nuphar lutea (Foti et al. 1994); Nymphaea odorata herbaceous alliance (Hoagland 1998a); Spatterdock - water lily wetland (Fike 1999); Natural Pond, in part (Smith 1991); Artificial Pond, in part (Smith 1991); Stable Natural Pool, in part (Smith 1991)

References: Ambrose 1990a, FNAI 1990, Faber-Langendoen et al. 1996, Fike 1999, Foti et al. 1994, Harris et al. 1996, Heineke 1987, Hoagland 1998a, Kovalchik 1993, Marr et al. 1980, Penfound 1952, Ramaley 1909, Sawyer and Keeler-Wolf 1995, Schafale and Weakley 1990, Smith 1991, Wharton 1978, Wolfe 1990

Authors: M. PYNE, MOD. M.S. REID, MP, Southeast Identifier: A.1984


NUPHAR LUTEA SSP. ADVENA - NYMPHAEA ODORATA HERBACEOUS VEGETATION
Broadleaf Pond-lily - White Water-lily Herbaceous Vegetation
Water-lily Aquatic Wetland                                                                                   G4G5 (96-10-03)
Ecological Group (SCS;MCS):                       
Eastern Open Ponds and Marshes (480-10; 1.4.1.1)

Concept: This rooted aquatic or open marsh community occupies shallow water depressions, oxbow ponds, backwater sloughs of river floodplains, slow moving streams, ponds, and small lakes throughout the central and eastern United States. It is dominated by rooted, floating-leaved aquatic species, with both submergent and emergent aquatics also present. Nuphar lutea ssp. advena and Nymphaea odorata are dominants. Other species present may include Brasenia schreberi, various Potamogeton spp., Polygonum amphibium, and Polygonum amphibium var. emersum (= Polygonum coccineum). Submerged aquatics more common in the southern part of the range include Cabomba caroliniana, Ceratophyllum demersum, and Heteranthera dubia.

Comments: Occurs in borrow pits on Kisatchie National Forest. On the Conecuh National Forest (Alabama), vegetation of this alliance occurs in Gum Pond and Open Pond as a mix of Nymphaea odorata and Nuphar lutea ssp. advena.

Range: This rooted aquatic community occupies shallow, quiet waters throughout the central and eastern United States, extending from Maine to Ontario and Minnesota, south to Oklahoma and east to Georgia.

States/Provinces: AL:S?, AR:S?, CT:S?, DE:S?, GA:S?, IA:SU, IL:S?, IN:S?, KY:S?, LA:S?, MA:S4, MD:S?, ME:S5, MI:S?, MN:S?, MO:S?, MS:S?, NC:S2, NH:S?, NJ:S4, NY:S5, OH:S?, OK:S?,S?, ON:S?, PA:S?, RI:S?, SC:S?, TN:S?, TX:S?, VA:S?, VT:S4, WI:S?, WV:S?

TNC Ecoregions: 31:C, 32:P, 36:C, 37:C, 39:C, 40:P, 41:C, 42:C, 43:C, 44:C, 45:C, 46:C, 47:C, 48:C, 49:C, 50:C, 51:C, 52:C, 53:C, 55:P, 56:C, 57:C, 58:C, 59:C, 60:C, 61:C, 62:C, 63:C

USFS Ecoregions: 212Aa:CCC, 212Ba:CCC, 212Bb:CCC, 212Ca:CCC, 212Cb:CCC, 212Da:CCC, 212Db:CCP, 212Dc:CCC, 212Fa:CCP, 212Fb:CCP, 212Fc:CCP, 212Fd:CCP, 212Ga:CCP, 212Gb:CCP, 212Hb:CPP, 212Ja:CCP, 212Jb:CCP, 212Jc:CCP, 212Je:CCP, 212Jf:CCP, 212Jj:CCP, 212Jl:CCP, 212Jm:CCC, 212Ka:CPP, 221Ah:CCC, 221Ai:CCC, 221Al:CCC, 221Am:CCC, 221Bd:CCP, 221Da:CCC, 221Db:CCC, 221Ea:CCC, 221Ed:CC?, 221Ef:CCC, 221He:CCC, 222Ch:CCC, 222Db:CCC, 222Gc:C??, 222Ha:CCC, 222Ja:CCC, 222Jb:CCC, 222Ji:CCC, 222Jj:CCC, 222Kf:CCC, 222Kg:CCC, 222Kh:CCC, 222Kj:CCC, 231Bc:CCC, 231Ga:CCC, 231Gb:CCC, 231Gc:CCC, 232Bf:CCC, 232C:CC, 232D:CC, 234Ac:CC?, 234An:CCC, 251Cf:CCC, 251Dd:CCC, 251Dg:CCC, 251Eb:CCC, M212Af:CCC, M212Bc:CCC, M212Bd:CCC, M212Cb:CCC, M212Cc:CCC, M212Ea:CCP, M212Eb:CCP, M221Aa:CCC, M221Ab:CCC, M221Ac:CCC, M221Ad:CCC, M221Ba:CCC, M221Bb:CCC, M221Bc:CCC, M221Bd:CCC, M221Be:CCC, M221Bf:CCC, M221Da:CCC, M221Dc:CCC

Federal Lands: DOD (Fort Benning); NPS (Acadia, Carl Sandburg Home); USFS (Angelina, Conecuh, Davy Crockett, Kisatchie, Ocala, Ozark, Sabine NF, Sam Houston?, Talladega); USFWS (Reelfoot)

Synonymy: Open Water/Aquatic Bed Veg., Natural Impoundment Pond (Ambrose 1990a) B, New England coastal plain pondshore (Rawinski 1984), L5D2aI1a. Nuphar lutea (Foti et al. 1994), Open water marsh with floating-leaved plants (NAP pers. comm. 1998)

References: Ambrose 1990a, Anderson 1982, Breden et al. 2001, FNAI 1990, Fike 1999, Fleming et al. 2001, Foti et al. 1994, Gawler 2002, Hoagland 2000, NAP pers. comm. 1998, NatureServe Ecology - Southeastern U.S. unpubl. data, Peet et al. 2002, Penfound 1953, Rawinski 1984, Schafale and Weakley 1990, Swain and Kearsley 2001, Zanoni et al. 1979

Authors: D. Faber-Langendoen, MCS Confidence: 3 Identifier: CEGL002386


NYMPHAEA ODORATA - ELEOCHARIS ROBBINSII HERBACEOUS VEGETATION
White Water-lily - Robbins Spikerush Herbaceous Vegetation
Coastal Plain Pond                                                                                                
G2 (98-12-07)
Ecological Group (SCS;MCS): 
Southeastern Coastal Plain Open Ponds and Marshes (345-05; n/a)



Concept: This hydromorphic Coastal Plain pond community ranges from southern New England to Maryland. It occurs in standing water in all but exceptionally dry years. The substrate is most often deep muck, but in oligotrophic ponds an organic layer may be absent or much reduced and the vegetation occurs on sand or mucky sand. Characteristic species include Nymphaea odorata, Nymphoides cordata, and Eleocharis robbinsii, with frequent associates including Scleria reticularis, Gratiola aurea, Proserpinaca pectinata, Utricularia juncea, Brasenia schreberi, Pontederia cordata, Ludwigia spp., Utricularia spp., and Eriocaulon aquaticum.

Range: This association is limited to the Atlantic and Coastal plains from southern New England to Maryland.

States/Provinces: DE:S?, MA:S2, MD:S?, NH:S?, NJ:S1S2, NY:S?, RI:S?

TNC Ecoregions: 58:C, 61:C, 62:C

USFS Ecoregions: 221A:CC, 221B:CC, 232Ac:CCC, 232C:CP

Synonymy: Coastal Plain Intermittent Pond (Breden 1989) B, New England coastal plain pondshore (Rawinski 1984)

References: Breden 1989, Breden et al. 2001, Rawinski 1984, Sneddon 1994, Swain and Kearsley 2001

Authors: ECS Confidence: 1 Identifier: CEGL006086

- Maryland Vegetation Classification Subset Report V.C. Hydromorphic rooted vegetation

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